Sanders McNew had a thread going that though not the same result I believe it to be the same cause.
Insufficient agitation in the initial stages of development.
Uneven development over flat skys or grey backgrounds*in his case*.
I don't think this is a exposure problem or camera problem.
Increase agitation and rotation of the film in the first 15 seconds of development.
To test for even development photograph a nuetral grey card and work on your agitation and rotation methods. I belive the problem will solved.