I thought those undercut rollers looked familiar!
All kidding aside, banding can also come from air pump pulsations in the drying cabinets, and pulsations in the emulsion feed pump. These all can cause banding in the final coating.
I have coated Titanox in gelatin on film many times. It is what we at Kodak called a "White Rug". It is very hard to coat uniformly at the level needed to get a good reflection backing for film or paper. I have 2 or 3 1 foot snips from a roll of one of my coatings sitting here in an old box. They sure have streaks in them. Making the TiO2 in gelatin is also not easy. It is tough to do and can require a ball mill.
This technology was used in Kodak Instant materials and in the Kodak Blood Test equipment now made by Johnson and Johnson at Kodak Park on their new updated equipment. They still have some old Kodak equipment running to make stuff for them too. So, the "White Rug" is still a coating staple at Kodak Park but made by J&J.